Phillip Glasser .... Fievel Mousekewitz (voice)
James Stewart .... Wylie Burp (voice)
Erica Yohn .... Mama Mousekewitz (voice)
Cathy Cavadini .... Tanya Mousekewitz (voice)
Nehemiah Persoff .... Papa Mousekewitz (voice)
Dom DeLuise .... Tiger (voice)
Amy Irving .... Miss Kitty (voice)
John Cleese .... Cat R. Waul (voice)
Jon Lovitz .... T.R. Chula (voice)
Jack Angel .... Additional Voices (voice)
Mickie McGowan .... Additional Voices (voice)
Fausto Bara .... Additional Voices (voice)
Larry Moss .... Additional Voices (voice)
Vanna Bonta .... Additional Voices (voice)
Nigel Pegram .... Additional Voices (voice)
Philip L. Clarke .... Additional Voices (voice) (as Philip Clarke)
Patrick Pinney .... Additional Voices (voice)
Jennifer Darling .... Additional Voices (voice)
Lisa Raggio .... Additional Voices (voice)
Annie Holliday .... Additional Voices (voice)
Lawrence Steffan .... Additional Voices (voice)
Sherry Lynn .... Additional Voices (voice)
David Tate .... Additional Voices (voice)
Lev Mailer .... Additional Voices (voice)
Robert Watts .... Additional Voices (voice)

Some time after the Mousekewitz's have settled in America, they find that they are still having problems with the threat of cats. That makes them eager to try another home out in the west, where they are promised that mice and cats live in peace. Unfortunately, the one making this claim is an oily con artist named Cat R. Waul who is intent on his own sinister plan. Unaware of this, the Mousekewitz's begin their journey west, while their true cat friend, Tiger, follows intent on following his girlfriend gone in the same direction.

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